B.S., Chemistry, Roanoke College, Salem, Virginia

Ph.D., Biochemistry, Duke University, Durham, North Carolina

Honors (Selected Academic and Professional):

American Heart Association Established Investigator, 1968-1973

Visiting Professor, Japan Society for Promotion of Science, 1978

National Heart, Lung, and Blood Institute MERIT Awardee, 1988-1997

Recipient, Federation of American Societies for Experimental Biology (FASEB) Excellence In Science Award, 1992

Wellcome Visiting Professor in the Basic Medical Sciences, 1992-1993

Member, U.S. National Committee of the National Academy of Sciences/ National Research Council to the International Union of Biochemistry & Molecular Biology, 1992-2003; Chair, 1997-2000.

Member, Advisory Panel on Biomedical Research of the Association of American Medical Colleges, 1995-1998

Elected, Institute of Medicine, National Academy of Sciences, 1996

Recipient, Bernard B. Brodie Award, American Society for Pharmacology and Experimental Therapeutics, 2000.

Elected, Fellow, American Association for the Advancement of Science, 2001.

Elected, Vice-President for Science Policy of the FASEB, 2001-2002.

Elected, President of the American Society for Biochemistry and Molecular Biology, 2000-2004.

Recipient, Doctorem Mediciniae (M.D.) Honoris Causae, Charles University, Prague, Czech Republic, 2005.

Public Advisory Committees (Selected):

Member, Board of Scientific Counselors, National Institutes of Environmental Health Sciences, 7/1/1982-6/30/1986; Chairman, 7/1/1984-6/30/1986

Member, Council for Extramural Grants of the American Cancer Society, 1998-2000.

Member, Advisory Committee to the Director of the National Institutes of Health, 12/1/2000-12/31/2004.

Member, National Advisory Research Resources Council, National Institutes of Health, 2005-2009.